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The shallower parts of the sea prevent migration of the deep water forms and no doubt living as they do in eternal darkness and in a temperature near the freezing point, there is little to induce them to much activity. The fact that they are easily captured in nets of comparatively small size would indicate that they move about slowly.
Dr. Jenkins, who has lately studied the fishes of the Sandwich Islands, informs me that less than five per cent. are found on our American coast, while a large per cent. is found all the way to the Red sea.
In other words, the fishes of the Sandwich Islands are East Indian rather than American. This is no doubt caused from the fact that the deep water between the islands of the American coast forms a barrier which has always prevented the two fish faunas from mingling with each other. Between Africa and the Sandwich Islands this has not been the case. A recent study of the fishes of the Galapagos Archipelago shows its fauna to be American, though in what respect its fishes differ from those of our west coast they resemble all the more the fishes of the Sandwich Islands.
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Two fish faunas will usually differ from each other if separated by an impassable barrier; especially is this true if the barrier be older than the two faunas.
Any barrier which prevents or hinders fishes in their movements from one body of water to another will separate two more or less well-marked fish faunas. These barriers may be mountains, or shallow water, as in the case of deep sea fishes; deep water, as in case of shore fishes; muddy or alkaline water, or water of different temperature. Temperature no doubt has far more influence in governing the movement of fishes than is generally believed. It plays an important part in guiding salmon up stream to their spawning beds. It explains why they reach the head waters of some streams and spawn earlier than in similar streams not far distant, but of different temperature. If you would know to what extent fishes of one region differ from those of another, study well the barriers between the two regions, learn to what extent and how long, they have existed, consider the age geologically of the two regions, and how fishes may have migrated to one or the other, and in a general way you will have the key to the sitnation, which a careful study of the fishes is quite sure to verify.
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